As for the main, it's the SMALLEST opening in the combined jet needle/main jet circuit that controls fuel flow.
The fuel first enters the main but the space between the TAPERED jet needle and needle jet is smaller at any throttle position less than 3/4. As the needle rises up through the needle jet, the thicker portion actually leaves the jet needle area and the lower/thinner portion comes into effect more and more. As the needle rises, the space between the two parts becomes larger than the opening of the main jet. At that point the main now controls flow because it is the smallest opening in the entire circuit.
As you can see, the tapered needle is designed to make a smooth/gradual transition to the main jet.


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